FOSSIL BATS OF THE AMERICAS
Eumops cf. E. perotis
McRae Wash, San Pedro Valley, Arizona, USA; Pliocene (Blancan); St. David Formation (Czaplewski 1993a).
Eumops sp. [image ]
Duke locality CVP 9, northeast of Villavieja, Magdalena River Valley, Huila Department, Colombia; middle Miocene (Laventan); Fish Bed, Villavieja Formation (Czaplewski 1997).
Chepe site, near Cuzco, northeast of Villavieja, Magdalena River Valley, Huila Department, Colombia; middle Miocene (Laventan); Las Lajas Member, Villavieja Formation (Czaplewski, Takai, et al. 2003c).
Type Specimen: Museo Geológico, Instituto Nacional de Investigaciones en Geosciencias, Minería y Quimica (INGEOMINAS), Bogotá, IGM 250997, right M1. [image ]
Type Locality and Horizon: Duke University locality CVP 9, northeast of Villavieja, Magdalena River Valley, Huila Department, Colombia; middle Miocene (Laventan); Fish Bed, Villavieja Formation.
Diagnosis (from Czaplewski 1997): M1 and M2 with hypocone tall, conical, and isolated from the postprotocrista; with trigon basin (=protofossa) that is closed posteriorly by the postprotocrista; with strong lingual cingulum; and with prominent paraconule and metaconule cristae that connect like keels to the bases of the paracone and metacone, respectively. M3 with strong paraconule crista and rudimentary hypocone.
Type Specimen: Museu Nacional de Rio de Janeiro 3000-V, partial skeleton with right P3, M1-M2, partial M3, partial right and left dentaries with p3, m1-m3.
Type Locality and Horizon: São Paulo state, Brazil; Oligocene (Deseadan); bituminous shales of Tremembé-Taubaté Basin (originally described by Paula Couto 1956 as Pleistocene; later revised by him and others to Oligocene).
Diagnosis (from Paula Couto 1956): Plus grande que les espèces vivantes sud américaines. M1-3 normales, mais portion interne légèrement comprimée antéro-postérieurement. Hypocône fort, bien détaché, sans bourrelet basal, beaucoup plus bas que le protocône. Protocône sans bourrelet basal. Parastyle, mésostyle et métastyle beaucoup plus bas que les cuspides principales. Mandibule forte, avec apophyse coronoïde élevée, apophyse angulaire forte et longue, dirigée vers larrière et vers le bas, et fosse massétèrique profonde. p4 et m1-3 normales. Paraconide plus petit et plus bas que le métaconide.
Remarks: The species was originally placed in Tadarida by Paula Couto 1956. It was moved to Mormopterus by Legendre 1984, who also erected a new subgenus (Neomops) for it.
Primary references: Paula Couto 1956; Legendre 1984a
Mormopterus or Tadarida
Thomas Farm local fauna, NE of Bell, Gilchrist County, Florida, USA; early Miocene (early Hemingfordian); Miocene clays and sands infilling a paleokarst sinkhole developed within Eocene marine limestone (two different species; Czaplewski et al. 2003b). [image ]
Type Specimen: Museo Geológico, Instituto Nacional de Investigaciones en Geociencias, Minería y Quimica (INGEOMINAS), Bogotá, IGM 184349, skull fragment including left maxilla with P4-M3, anterior root of zygomatic arch, anterior margin of orbit, and portions of the palate, plus associated radius fragment.
Type Locality and Horizon: Duke locality 90, northeast of Villavieja, Magdalena River Valley, Huila Department, Colombia; middle Miocene (Laventan); about 60 m above Chunchullo Sandstone Beds, La Victoria Formation.
Diagnosis (from Czaplewski 1997): A large molossid, similar in the size of its teeth to Eumops perotis (smaller than E. dabbenei; larger than Cheiromeles parvidens but smaller than C. torquatus, the largest living molossid bat), and unique in that its upper first and second molars lack a hypocone and lingual cingulum but have a postprotocrista that is continuous with the cingulum of a small talon, which, in turn, is continuous with the postcingulum.
Type Specimen: Harvard University, Museum of Comparative Zoology 49076, cranium.
Type Locality and Horizon: New Cave (now known as Slaughter Canyon Cave), Carlsbad Caverns National Park, Eddy County, New Mexico, USA; Pleistocene; cave deposits.
Diagnosis (from Lawrence 1960): This form is characterized by its relatively great size and long, rather evenly rectangular skull.
Primary references: Lawrence 1960; Morgan, in press.
Macasphalt Shell Pit, Sarasota County, Florida, USA; late Pliocene (late Blancan); Pinecrest Sand Member, Tamiami Formation (Morgan and Ridgway 1987; Morgan 1991; Czaplewski et al. 2003b). [image ]
Remarks: A specimen originally reported as bat, near Tadarida by Dalquest 1975 in the Pliocene Blanco local fauna of Texas, USA, is actually an indeterminate vespertilionid (Czaplewski et al. 2003b).
Type Specimen: Royal Saskatchewan Museum SMNH P1654.311, left M1.
Type Locality and Horizon: Swift Current Creek, Saskatchewan, Canada; middle Eocene (late Uintan), Cypress Hills Formation.
Diagnosis (from Storer 1984, who considered Wallia to be a proscalopid insectivoran): Molars strongly dilambdodont; upper molars with oblique postprotocrista, indistinct para- and metaconule, large, low hypocone except in M3, strong preprotocrista forming anterior shelf reaching to strongly inflected parastyle; lower molars with anteroposteriorly compressed, oblique trigonid and talonid, continuous, strong anterior, buccal and posterior cingula, hypoconulid as high as postcristid.
Remarks: Legendre 1985 referred Wallia to the Molossidae.
Primary references: Storer 1984; Legendre 1985
Molossidae, genus and species indet.
Brooksville 2 locality, Hernando County, Florida, USA; late Oligocene (early Arikareean); clay and sand infilling of karstic solution cavities developed within the marine, early Oligocene Suwannee Limestone (Czaplewski et al. 2003b). [image ]